黄土高原山地森林群落植物区系特征与地理格局

doi:10.11821/dlxb201407004

Abstract

Abstract:

Based on the sampling data, we analyzed and revealed the floristic characteristics and the relationships of floristic composition between and among 11 mountains on the Loess Plateau. The results indicate that: (1) The floristic composition in forest communities in this region is abundant. Among the 353 plots, 1222 species of vascular plants, belonging to 473 genera of 108 families, were discovered. In those species, 1179 species of 447 genera of 93 families are Angyospermae, 9 species of 7 genera of 4 families are Gymnospermae, and 34 species of 19 genera of 11 families are Pteridophyta. (2) On the regional and local scales, the dominant distribution types of floristic elements at the family and genus levels are temperate origin, especially the north temperate origin. This characteristic is unique on the scale of community level. (3) The first axis of NMDS ordination revealed typical variation range of elevational gradient. The second axis displayed the latitudinal relationship between and among different mountains. The whole study area can be divided into 4 groups with different levels of floristic linkage by hierarchical clustering. Through the comparison of maximum Sørenson similarity coefficient, Helan Mountain, located at the northwest edge of the region, had the weakest relation with other mountains, whereas Taiyue Mountain had the strongest relation with all of the other mountains. (4) Multiple linear regression analysis accounted for 42.1% of the variation within the R/T value (ratio of the tropical to the temperate elements), revealing a significant trend that the R/T value in the region increased at sites with high annual average temperature, whereas it decreased at sites with high elevation. However, no significant tendency of the R/T value with annual precipitation was found in the same analysis.

Key words: Loess Plateau, forest community, plantflora, R/T value, geographic pattern

Mingfei ZHAO, Yuhang WANG, Kaixiong XING, Muyi KANG, Quanru LIU, Qiuyi LI, Yongmei HUANG. Floristic traits and geographic patterns of natural montane forests at community level on the Loess Plateau, North China[J].Acta Geographica Sinica, 2014, 69(7): 916-925.

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References 36

[1]	Wu Zhengyi, Wang Hesheng.Natural Geography of China: Plant Geography (1st volume). Beijing: Science Press, 1983.
	[吴征镒, 王荷生. 中国自然地理: 植物地理 (上册). 北京: 科学出版社, 1983.]
[2]	Wu Zhengyi, Zhou Zhekun, Sun Hang et al. The Areal Types of Seed Plants and Their Origin and Differentiation. Kunming: Yunnan Science and Technology Press, 2006.
	[吴征镒, 周浙昆, 孙航等. 种子植物分布区类型及其起源和分化. 昆明: 云南科技出版社, 2006.]
[3]	Zhu Fenmeng, An Shuqing, Guan Baohua et al. A review of ecotone concepts, attributes, theories and research advances. Acta Ecologica Sinica, 2007, 27(7): 3032-3042.
	[朱芬萌, 安树青, 关保华等. 生态交错带及其研究进展. 生态学报, 2007, 27(7): 3032-3042.]
[4]	Wang Hesheng.The evolution and sources of north China's flora. Acta Geographica Sinica, 1999, 54(3): 213-223.
	[王荷生. 华北植物区系的演变和来源. 地理学报, 1999, 54(3): 213-223.]
[5]	Di Weizhong, Tian Lianshu, Li Zhijun.The characteristics of Helan Mountain flora of spermatophytes. Acta Botanica Boreali-Occidentalia Sinica, 1988, 8(4): 242-254.
	[狄维忠, 田连恕, 李智军. 贺兰山种子植物区系的特征. 西北植物学报, 1988, 8(4): 242-254.]
[6]	Chai Baofeng, Li Cui.The characteristics of spermatophytes in Taiyue Mountains of Shanxi Province. Mountain Research, 1997, 15(3): 162-166.
	[柴宝峰, 李毳. 山西太岳山种子植物区系特征. 山地研究, 1997, 15(3): 162-166.]
[7]	Zhang Feng, Shangguan Tieliang.On the flora of seed plants in Guandi Mountains, Shanxi. Bulletin of Botanical Research, 1998, 18(1): 20-27.
	[张峰, 上官铁梁. 山西关帝山种子植物区系研究. 植物研究, 1998, 18(1): 20-27.]
[8]	Shangguan Tieliang, Zhang Feng, Qiu Fucai.Study on flora diversity of the seed plants in Luya Mountain Nature Reserve, Shanxi. Journal of Wuhan Botanical Research, 1999, 17(4): 323-331.
	[上官铁梁, 张峰, 邱富财. 芦芽山自然保护区种子植物区系地理成分分析. 武汉植物学研究, 1999, 17(4): 323-331.]
[9]	Dai Junhu, Bai Jie, Shao Liyang et al. A primary analysis of the flora on Liupan Mountains: Its nature and characteristics. Geographical Research, 2007, 26(1): 91-100.
	[戴君虎, 白洁, 邵力阳等. 六盘山植物区系基本特征的初步分析. 地理研究, 2007, 26(1): 91-100.]
[10]	Wang Tiejuan, Zhao Yizhi, Wang Nana et al. Study on the flora of seed plant in Daqing Mountain. Journal of Arid Land Resources and Environment, 2008, 22(1): 174-178.
	[王铁娟, 赵一之, 王娜娜等. 内蒙古大青山种子植物区系研究. 干旱区资源与环境, 2008, 22(1): 174-178.]
[11]	Ling Dengwu.The floristic geography of vascular plant in the Loess Plateau of northern Shaanxi Province [D]. Northwest Agriculture and Forestry University, 2008.
	[李登武. 陕北黄土高原维管植物区系地理研究[D]. 西北农林科技大学, 2008.]
[12]	Zhang Xibiao.Floristic characteristics of seed plants in Ziwuling of Loess Plateau. Chinese Journal of Ecology, 2005, 24(8): 872-877.
	[张希彪. 黄土高原子午岭种子植物区系特征研究. 生态学杂志, 2005, 24(8): 872-877.]
[13]	Shangguan Tieliang, Zhang Feng.Analysis on the flora of xylophyta in Zhongtiao Mountains, Shanxi. Bulletin of Botanical Research, 2000, 20(2): 143-155.
	[上官铁梁, 张峰. 中条山木本植物区系地理成分分析. 植物研究, 2000, 20(2): 143-155.]
[14]	Lian Kaimin, Zhang Li, Zhao Lulu et al. Analysis on the floristic characteristics of wild seed plants in the Zhongtiao Mountains, Shanxi. Journal of Wuhan Botanical Research, 2010, 28(2): 144-152.
	[廉凯敏, 张丽, 赵璐璐等. 山西中条山野生种子植物区系研究. 武汉植物学研究, 2010, 28(2): 144-152.]
[15]	Yang Qinye, Yuan Baoyin.Natural Environment and Its Evolution in Loess Plateau. Beijing: Science Press, 1991.
	[杨勤业, 袁宝印. 黄土高原地区自然环境及其演变. 北京: 科学出版社, 1991.]
[16]	Wu Qinxiao, Yang Wenzhi.Vegetation construction and sustainable development on the Loess Plateau. Beijing: Science Press, 1998.
	[吴钦孝, 杨文治. 黄土高原植被建设与持续发展. 北京: 科学出版社, 1998.]
[17]	Li Bin.Analysis of relationships between vegetation and climate variables in Loess Plateau. Acta Ecologica Sinica, 2003, 23(1): 82-89.
	[李斌. 黄土高原地区植被与气候的关系. 生态学报, 2003, 23(1): 82-89.]
[18]	Fang Jingyun, Wang Xiangping.Methods and protocols for plant community inventory. Biodiversity Science, 2009, 17(6): 533-548.
	[方精云, 王襄平. 植物群落清查的主要内容, 方法和技术规范. 生物多样性, 2009, 17(6): 533-548.]
[19]	Ma Keping, Gao Xianming.On the characteristics of the flora of Dongling Mountain area and its relationship with a numberof other mountains floras in China. Bulletin of Botanical Research, 1995, 15(4): 501-515.
	[马克平, 高贤明. 东灵山地区植物区系的基本特征与若干山区植物区系的关系. 植物研究, 1995, 15(4): 501-515.]
[20]	Faith D P.Compositional dissimilarity as a robust measure of ecological distance. Vegetatio, 1987, 69(1-3): 57-68.
[21]	Stallins J A.Dune plant species diversity and function in two barrier island biogeomorphic systems. Plant Ecology, 2003, 165(2): 183-196.
[22]	Zhang Yili, Zhang Xuemei.Coefficient of similarity: An important parameter in floristic geography. Geographical Research, 1998, 17(4): 94-99.
	[张镱锂, 张雪梅. 植物区系地理研究中的重要参数——相似性系数. 地理研究, 1998, 17(4): 94-99.]
[23]	Peng Hua.The floristic equilibrium point of seed plants in Mt. Wuliangshan. Acta Botanica Yunnanica, 1996, 18(4): 385-397.
	[彭华. 无量山种子植物的区系平衡点. 云南植物研究, 1996, 18(4): 385-397.]
[24]	Oksanen J, Kindt R, . Acesso em, 2007, 10(1): 2008.
[25]	Zhang Wenhui, Li Dengwu.The characteristics of the seed plant flora in Loess Plateau. Bulletin of Botanical Research, 2002, 22(3): 373-379.
	[张文辉, 李登武. 黄土高原地区种子植物区系特征. 植物研究, 2002, 22(3): 373-379.]
[26]	He Xiaoqun. Multivariate Statistical Analysis.Beijing: China Renmin University Press, 2008.
	[何晓群. 多元统计分析. 北京: 中国人民大学出版社, 2008.]
[27]	Wang Hesheng.Floristic Plant Geography in North China. Beijing: Science Press, 1997.
	[王荷生. 华北植物区系地理. 北京: 科学出版社, 1997.]
[28]	Wiens J J, Donoghue M J.Historical biogeography, ecology and species richness. Trends in Ecology and Evolution, 2004, 19(12): 639-644.
[29]	Götzenberger L, de Bello F, Bråthen K Aet al. Ecological assembly rules in plant communities: Approaches, patterns and prospects. Biological Reviews, 2012, 87(1): 111-127.
[30]	Feng Jianmeng, Dong Xiaodong, Xu Chengdong.Effects of sampling scale on latitudinal patterns of species diversity in seed plants in northwestern Yunnan, China. Biodiversity Science, 2009, 17(3): 266-271.
	[冯建孟, 董晓东, 徐成东. 取样尺度效应对滇西北地区种子植物物种多样性纬度分布格局的影响. 生物多样性, 2009, 17(3): 266-271.]
[31]	Zhang Na.Scale issues in ecology: Concepts of scale and scale analysis. Acta Ecologica Sinica, 2013, 26(7): 2340-2355.
	[张娜. 生态学中的尺度问题: 内涵与分析方法. 生态学报, 2013, 26(7): 2340-2355.]
[32]	Cox C B, Moore P D.Biogeography. Zhao Tieqiao trans. Beijing: Higher Education Press, 2007.
	[Cox C B, Moore P D.生物地理学. 赵铁桥译. 北京: 高等教育出版社, 2007.]
[33]	Feng Jianmeng, Xu Chengdong.Flora transition and its biogeographical significance. Chinese Journal of Ecology, 2009, 28(1): 108-112.
	[冯建孟, 徐成东. 植物区系过渡性及其生物地理意义. 生态学杂志, 2009, 28(1): 108-112.]
[34]	Fang jingyun. Re-discussion about the forest vegetation zonation in eastern China. Acta Botanica Sinica, 2001, 43(5): 522-533.
	[方精云. 也论我国东部植被带的划分. 植物学报, 2001, 43(5): 522-533.]
[35]	Currie D J, Francis A P.Regional versus climatic effect on taxon richness in angiosperms: Reply to Qian and Ricklefs. The American Naturalist, 2004, 163(5): 780-785.
[36]	Feng Jianmeng, Xiong Junling, Xu Chengdong.Geographical patterns of floristic transition of seed plant in north-south range-gorge region in western Yunnan and the biogeographical significance. Acta Botanica Boreali-Occidentalia Sinica, 2009, 29(2): 373-378.
	[冯建孟, 熊珺玲, 徐成东. 云南西部纵向岭谷区种子植物区系过渡性的地理分布格局及其生物地理意义. 西北植物学报, 2009, 29(2): 373-378.]

山地名称	海拔 (m)	调查范围 (海拔m)	调查涉及的典型群落建群种	样地数
贺兰山	3556	1330~3150	青海云杉 (Picea crassifolia)、油松 (Pinus tabulaeformis)、灰榆 (Ulmus glaucescens)、杜松 (Juniperus rigida)	91 (其中65个为 10 m×10 m样地)
大青山	2338	1150~1940	白桦 (Betula platyphylla)、山杨 (Populus davidiana)、油松	15
芦芽山	2787	1820~2640	华北落叶松 (Larix principis-rupprechtii)、青杄 (Picea wilsonii)、白桦	17
关帝山	2831	1700~2510	华北落叶松、青杄、白桦、辽东栎 (Quercus wutaishanica)、油松	22
太岳山	2354	1480~2220	华北落叶松、辽东栎、红桦 (Betula albo-sinensis)、白桦、油松、青榨槭 (Acer davidii)	17
中太行	2148	1510~1940	辽东栎、山杨、油松	9
中条山	2322	730~2280	油松、辽东栎、栓皮栎、鹅耳枥 (Carpinus turczaninowii)、千金榆 (Carpinus cordata)、葛萝槭 (Acer grosseri)、栓皮栎 (Quercus variabilis)、槲栎 (Quercus aliena)、华山松 (Pinus armandii)	83 (20 m×20 m样地)
黄龙山	1784	1320~1740	辽东栎、油松、白桦	11
子午岭	1758	1120~1730	辽东栎、油松、白桦、侧柏 (Platycladus orientalis)	45
关山	2200	1520~2090	白桦、辽东栎、油松、槲栎、胡桃楸 (Juglans mandshurica)	18
六盘山	2942	2110~2710	白桦、红桦、糙皮桦 (Betula utilis)、辽东栎	22

分布区类型	科水平		属水平
分布区类型	No.	(%)	No.	(%)
F1. 世界广布Cosmopolitan	42	-	57	-
F2. 泛热带分布及变型Pantropic	27	41.5	31	7.5
F3. 热带亚洲和热带美洲间断分布Trop. Asia & Trop. Amer. Disjuncted	4	6.2	5	1.2
F4. 旧世界热带分布及变型Old World Trop.	1	1.5	9	2.2
F5. 热带亚洲至热带大洋洲分布Trop. Asia to Trop. Australasia Oceania	1	1.5	5	1.2
F6. 热带亚洲至热带非洲分布Trop. Asia to Trop. Africa	-	-	5	1.2
F7. 热带亚洲分布及变型Trop. Asia	1	1.5	8	1.9
F8. 北温带分布及变型North Temperate	23	35.4	167	40.1
F9. 东亚和北美间断分布E. Asia & N. Amer. Disjuncted	2	3.1	40	9.6
F10. 旧世界温带分布Old World Temp.	1	1.5	67	16.1
F11. 温带亚洲分布Temp. Asia	-	-	24	5.8
F12. 地中海区、西亚至中亚分布Med., W. Asia to C. Asia	-	-	7	1.7
F13. 中亚分布C. Asia	-	-	2	0.5
F14. 东亚分布E. Asia	6	9.2	37	8.9
F15. 中国特有分布Endemic to China	-	-	9	2.2

生态因子	回归系数	标准化回归系数	t值	p值	决定系数	F值	p值
海拔 (m)	-0.00049	-0.2399	-3.265	0.001	0.421	59.41	< 0.001
年均温 (°C)	0.10170	0.2904	4.419	< 0.001
年降水 (mm)	-0.00002	-0.004	-0.082	0.935

Floristic traits and geographic patterns of natural montane forests at community level on the Loess Plateau, North China

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